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actriia actriib alk2 alk3 alk6 antagonist arterial bambi betaglycan binding bone brachydactyly bram1 brii caronte caveolin cerberus cgk1 chordin chrd c-kit coco contact crossveinless c-src cytosolic dan dante dorsalizing dragon dullard email endoglin eps15r fibrodysplasio fop fusion gene gremlin healing history hypertension inhibition inhibitory interaction i-smad knockout ligand limk1 noggin nonfusion organogenesis ossificans osteoporosis pah pp2a prdc processing progressiva pulmonary rack1 rgma rgmb rgmc ror2 r-smad sclerostin sf3b4 signaling spinal spine stimulatory tctex1 tgf-beta transduction trb3 trkc tsg usag1 ventralizing xiap